I.  WHAT IS GENETIC DRIFT?

     A.  Definition

     B.  Analogy--Drawing marbles from an urn

     C.  Long-term consequences of genetic drift: Computer simulation of genetic drift

     D.  Properties of genetic drift  (see Handout)
 

II.   Empirical analysis of genetic drift: Buri's studies on drift in lab populations of
   D. melanogaster

     A.  Behavior of ensembles of populations under genetic drift

     B.  Buri's Experiment: Overview

     C.  Preliminary experiments

         1.  (see Measuring Fitness in Drosophila  laboratory populations).

         2.  Results

     D.  Main experiment

        1.  Procedure

        2.  Results

     E.  Conclusions: Genetic drift in these laboratory populations behaves according to theory.